Puppies do feel ashamed when they have done something wrong and are reprimanded for it; you find them whining alone at a corner. When he does a wrong thing and you give him strong eye contact and reproof in a firm tone, he learns to associate the act with a wrong and avoids the mistake.
Animals feel whole range of emotions n feelings superbly more than humans, but because humans cannot comprehend. Humans believe they r falsely superior, since human values r physical n sighted. While animals senses beyond humans, generally they get along better with each other meow